INTERMEDIATE
DISTURBANCE HYPOTHESIS:
The Intermediate Disturbance Hypothesis states that diversity is
maximized
when ecological disturbance is neither too rare nor too frequent. At low levels
of disturbance, more competitive organisms will push subordinate species to
extinction and dominate the ecosystem. At
high levels of disturbance, due to frequent forest fires or human impacts like
deforestation, all species are at risk of going extinct(both r and k-selected
species). According to intermediate disturbance theory, at intermediate levels of disturbance,
diversity is thus maximized because both competitive k-selected selected and
opportunistic r-selected species can coexist
This coexistence is a result of the
differing life history strategies of species,which dictate a preference for high or low
disturbance. K-selected species tend to be more competitive, because they
invest a larger proportion of resources into growth and competition and thus
generally dominate stable ecosystems over long time periods. In contrast,
r-selected species, which colonize open areas quickly, can dominate landscapes
recently cleared by
disturbance.
Therefore the areas where disturbance occurs frequently, but in intermediate
magnitude both r-selected and k-selected species exist.
The intermediate disturbance hypothesis is
consistent with diversity patterns observed in natural and altered lotic ecosystems. Species diversity is reduced in stream habitats exposed to levels
of disturbances that are severe. In addition, habitats with enhanced
environmental consistency exhibit suppressed diversity even if adverse
conditions (e.g., oxygen deficits) are not apparent. It is postulated that
‘undisturbed’ lotic systems are in fact ‘disturbed’ and that the high biotic diversity of natural streams is a function of moderate perturbation. Diversity
is enhanced by the spatio-temporal heterogeneity resulting from intermediate
disturbance, which maintains the community in a non-equilibrium state (a state
where both ‘k’ as well as ‘r’-selected species co-exist.
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